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Fig. 1. Aggregate separation scheme. Fractionation I involved wet sieving air-dry soil into four aggregate-size classes. In fractionation II, small macroaggregates (250–2000 µm) were further separated into coarse sand and POM (>250 µm), microaggregates (53–250 µm) and the silt and clay fraction (<53 µm).

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